Species | Adults per breeding group a | Natal emigration b | Natal or nonnatal c | Transitional period d | Ultimate fate of emigrants e | Aggression to maturing natal individuals f | Resistance to immigrants g | References |
Pan troglodytes | ||||||||
| 7-25 | r | N | Ab | J; return to natal | Rare but potentially serious (F) | H (F); males may protect | Wrangham (1975)*; Goodall (1977) |
| 5-16 | R | N(?) | R; S | ? | V (M); peer rivals, coalitions important | ?; males hostile to foreign males | Nishida (1979)*; Pusey (1980)*; additional refs ²h |
Colobus badius tephrosceles | ||||||||
| 6-21 | O/r | N | R/Ab | J | L | L | Struhsaker (1975)* |
| 3-10 | O/r | N(?) | r; S | J; some permanent solitary? | Y (M); mortality inferred, some/many expelled | H (M) | Struhsaker & Leland (1979) |
Macaca radiata | ||||||||
| 4-18 (9.7) | O/r | ? | R (?) | J | L | ? | Simonds (1965)*; Sugiyama (1971)*; |
| 1-11 (8) | O/r cohorts | ? | R (?) | J; often cohort (?) | Y (M); subadult killed & wounds common | L | Wade (1979); Ali (1981) |
Papio hamadryas | ||||||||
| 1-10 (1.9) | Clan: r; Band: O | B | Ab | J, F | L | V (F); Fs may promote entry | Kummer (1968)*; Kummer (pers. comm.) |
| 1-2 (1 breed) | Clan: O, rarely permanent | Forms loose assoc. w/unit in same/diff. band; [cont -->] | eventual return & J/F in natal clan | ? | F transfer assoc. w/M-M fighting | Sigg et al. (1982) | |
Gorilla gorilla beringei | ||||||||
| 2-10 (6.2) | r/A | B | Ab/R | J, F | Rare but potentially serious (F) | L | Schaller (1963)*; Harcourt et al. (1976); |
| 1-4 (1.7) | r | N | r; S | F; return to natal | L but prevented from breeding | H; primarily by silverback | Harcourt (1978); Harcourt & Fossey (1981); Harcourt (pers. comm.) |
Presbytis senex | ||||||||
| 1-7 (4.1 & 3.3) | O | N | r; G | J (some F?) | By new M after takeover | ? | Rudran (1970, 1973)* |
| 1 (rare 2) | r/A | N/B | r; G | R (some F?) | As above & poss. by "old" male | H (M); probable deaths | Manley (1978) |
Presbytis johnii | ||||||||
| 1-5 (2.6) | O | N/B | r; S, G, or congeneric species | J (some F?) | L | L | McCann (1932); Poirier (1969)* |
| 1 (rare 2) | r/A | N/B | r; G | R (some F?) | Probable but not observed | V (M); none in Poirier (1969) | Oates et al. (1980); Moore (pers. obs.) |
Alouatta seniculus | ||||||||
| 1-5 (2.7) | r/O | N | r; S, G | J, F | V (F); may depend on troop size | H (F) | Rudran (1979)*; Gaulin & Gaulin (1982); |
| 1-3 (1.7) | r/A | B | r; S, G | R, F | ? | H (M); some deathsi | Sekulic (1982a, b) |
Alouatta palliata | ||||||||
| 4-16 (6) | r | N | r; S | J, F | Y (F); contributes to emigration | H (F) | Chivers (1969)*; Glander (1975a, b, 1980); |
| 2-6 (3.3) | r | N(?) | r; S | J, F, R | Y (M); may depend on group size & sex ratio | H (M & F) | Scott et al. (1978); Jones (1980)*; Estrada (1982) |
Colobus badius rufomitratus | ||||||||
| 7-18 (9.6) | r (?) | B | R | J | L | L | Marsh (1979a*, b) |
| 1-2 (1.5) | r/A | B | r; S or unstable G | R | Y (M) | V (M); from none to intense |
NOTES
aRange (mean); calculated from the reference marked with an asterisk. [eprint note: references are separated by sex only for format purposes; they are not sex-specific].
bNatal emigration: A, all; r, routine; O, occasional; R, rare.
cNatal or nonnatal: N, most emigration is natal; B, both natal and breeding emigration occurs (see Greenwood, 1980)
dTransitional period: Ab, absent; R, rare; r, routine. If there is a transitional period, the emigrant may spend it as a solitary (S) or in a nonbreeding group (G).
eFate of emigrants: J, join an existing group peacefully; F, form a new group; R, join an existing group by replacing a peer(s).
fAggression to maturing individuals: L, little if any; Y, normally yes; V, variable; (M/F), the sex that is responsible for most of the aggression.
gResistance to immigrants: L, little if any; H, high; V, variable; (M/F), the sex which resists
hAdditional references: Bygott (1979); Goodall et al. (1979); Sugiyama & Koman (1979); Nishida (1981); Sugiyama (1981).
iMale A. seniculus may accept or possibly encourage a second male if by doing so they can form a coalition that can resist takeover attempts by other males (Sekulic, personal communication, 1982c).